Itilochelys

Extinct genus of turtles

Itilochelys Temporal range: Danian PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Superfamily:	Chelonioidea
Family:	Cheloniidae
Genus:	†Itilochelys
Species:	†I. rasstrigin
Binomial name
†Itilochelys rasstrigin Danilov, Averianov, & Yarkov, 2010

Itilochelys is an extinct genus of sea turtle in the family Cheloniidae containing the single species Itilochelys rasstrigin. The species is known only from the Early Paleocene, Danian stage Rasstrigin 2 locality, Dubovsky District, Volgograd Oblast, Russia.^[1]

History and classification

Itilochelys rasstrigin is known only from three fossils, the holotype, specimen number "ZIN PH 1/118" and two referred specimens, numbers "ZIN PH 2/118" and "ZIN PH 3/118" . The type specimen is composed of a grouping of cervical vertebrae I-III, partial skull, and disarticulated lower jaw. Specimen ZIN PH 2/118 is an isolated section of dentary while ZIN PH 3/118 is a right humerus. The fossils were all recovered by A. A. Yarkov from fossiliferous outcrops of the Beryozovaya beds exposed at the Rasstrigin 2 locality.^[1] As of 2011, the type and referred specimens are preserved in the Paleoherptological collections housed at the Zoological Institute of the Russian Academy of Sciences, in Saint Petersburg, Russia. Itilochelys was first studied by I. G. Danilov and A. O. Averianov of the Zoological Institute along with A. A. Yarkov of the Natural History, Humanitarian Institute in Volgograd. Their 2010 type description of the genus and species was published in the journal Proceedings of the Zoological Institute RSA.^[1] The generic name was coined by Danilov, Averianov, and Yarkov as a combination of the medieval Arabian/Persian name of the Volga River, Itil, along with the Greek "chelys", meaning turtle. The specific epithet rasstrigin is in reference to the type locality of the specimen.^[1]

Itilochelys rasstrigin is one of three fossil cheloniid sea turtle taxa which have been recovered from the Volgograd region and the most complete specimen recovered so far. The other two taxa are represented by a specimen of the genus Euclastes known from the Karpovka site and a specimen related to the genus Tasbacka from the Malaya Ivanovka site.^[1]

The placement of Itilochelys rasstrigin in the family Cheloniidae is supported by the contact of the squamosal and parietal bones, an emarginated v-shaped crest on the underside of the basisphenoid, and features of the nasal bone structure. Placement of the species as a stem member in the family is based on the structure of the humerus which shows a morphology transitioning towards the derived pelagic structuring of modern cheloniids.^[1]

References

^ ^a ^b ^c ^d ^e ^f Danilov, I.G.; Averianov, A.O; Yarkov, A.A. (2010). "Itilochelys rasstrigin gen. et sp. nov., a new hard-shelled sea turtle (Cheloniidae sensu lato) from the Lower Palaeocene of Volgograd Province, Russia" (PDF). Proceedings of the Zoological Institute RAS. 314 (1): 24–41. doi:10.31610/trudyzin/2010.314.1.24. S2CID 250006012.

Cheloniidae family

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Subclass: Anapsida
Order: Testudines
Suborder: Cryptodira
Superfamily: Sea turtles
Family: Cheloniidae

Genera

Species of the Cheloniidae family

†Allopleuron

†Allopleuron hofmanni

†Argillochelys

†Argillochelys africana

Caretta

Loggerhead sea turtle

†Carolinochelys

†Carolinochelys wilsoni

Chelonia

Green sea turtle

†Eochelone

†Eochelone brabantica
†Eochelone voltregana

Eretmochelys

Hawksbill sea turtle

†Gigantatypus

†Gigantatypus salahi

†Glarichelys

†Glarichelys knorri

†Itilochelys

†Itilochelys rasstrigin

Lepidochelys

Kemp's ridley sea turtle
Olive ridley sea turtle
†Lepidochelys waikatoica

†Mexichelys

†Mexichelys coahuilaensis

†Miocaretta

†Miocaretta lankae

Natator

Flatback sea turtle

†Pacifichelys

†Pacifichelys hutchisoni
†Pacifichelys urbinai

†Syllomus

†Syllomus aegyptiacus
†Syllomus crispatus

†Tasbacka

†Tasbacka aldabergeni
†Tasbacka danica
†Tasbacka ouledabdounensis
†Tasbacka ruhoffi
†Tasbacka salisburgensis

Phylogenetic arrangement of turtles based on turtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. †=Extinct.

Testudines

Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Diapsida
Order: Testudines

Suborder

Superfamily

Family

Genus

Cryptodira

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based on Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.